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As Jak, Daxter, and Pecker lose consciousness, they are rescued by a group of desert-dwellers, who take them back to Spargus City. The Palace is then destroyed by an unknown force.

As they travel through the desert, flashbacks reveal that Haven City is at war between the Freedom League and the surviving Metal Heads and their allies, the Krimzon Guard Death Bots.

Ashelin, who opposes the banishment, gives Jak a beacon before she leaves and tells him to 'stay alive'. Story The game begins with and being banished into the Wasteland by the tyrannical Count Veger for supposed crimes against Haven City. The Dark Makers are Precursors who have been corrupted by Dark Eco, similar to. Even as he attempts this, a species known as the Dark Makers begin to invade the planet, seeking to destroy it. The player later finds that Veger attacked the Palace himself, wishing to journey to the core of the planet and gain the power of the Precursors. A new secondary villain is, a self-absorbed Haven City aristocrat who banishes Jak to the wasteland at the beginning of the game. Few of the original from reappear in this game. More minor but returning characters include Sig, a spy for Damas back in Haven City who later becomes the new king of Spargus after Damas is killed, Jinx, a former member of the Underground, Vin, whose brain now resides inside a computer and Pecker, Onin's translator from Jak 2 becomes an adviser to Damas in the early parts of the game. It is revealed that not only did Damas lose his son, but was the leader of Haven City before being betrayed by Baron Praxis and being banished to the desert.